Biosphere Contents

Glossary

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Abiotic
Environmental factors that are nonliving components of the ecosystem.

Acetic acid
An organic acid produced by certain micro-organisms during the decomposition of organic matter. Also known as ethanoic acid.

Acid
A substance which, on being dissolved in water produces hydrogen ions at concentrations which exceed those found in pure water. Acids are strongly corrosive and will dissolve metals from many natural materials, e.g. rock.

Active surface
The term active surface is used in the text to denote the general attributes of any form of surface.

Allogenic succession
Succession due to external factors. (See: Succession).

Amensalism
An ecological interaction where the competition is asymmetrical and the interaction is negative for only one participant ( - 0 interaction).

Amino acids
An organic compound containing an amino group and a carboxyl group; amino acids are the units or building blocks that make peptide and protein molecules.

Ammonification
Conversion of an organic form of nitrogen (e.g. of protein) into the inorganic ammonium (NH4+) ion.

Anaerobiosis
The state of being oxygen free. Describes the breakdown of sugars in organic compounds in the absence of oxygen (releasing some energy and producing organic acids and/or alcohol).

Aquatic ecosystem
Any ecosystem of which the principal component is water, such as ponds, lakes, rivers, streams and oceans. Lawrence (1995)

Asexual reproduction
Reproduction which forms new individuals from a single individual without the involvement of gametes. The offspring are genetically identical to the parent and to each other. Parker (1984) Lawrence (1995)

Assimilation
In autotrophic organisms it is the uptake of elements and simple inorganic compounds such as carbon dioxide, nitrogen and water from the environment and their incorporation into complex organic compounds. In heterotrophs it is the conversion of digested food material into complex biomolecules. Lawrence (1995)

Atmosphere
The gaseous envelope surrounding planets. The term is generally applied to the whole body of terrestrial air. The earth's atmosphere is composed mainly of nitrogen, oxygen, argon and carbon dioxide.

Autogenic succession
Succession occurring due to changes brought about by the presence of vegetation itself.

Bacteria
A group of extremely metabolically diverse, prokariotic, unicellular micro-organisms, usually possessing cell walls. They are ubiquitous in soil and water. The many forms include parasitic and disease-causing organisms and those which play an important part in the breakdown of dead organisms. Lawrence (1995)

Biodiversity
The diversity of the natural world, ecological variety and richness. Chambers (1994)

Biofilm
A thin film composed of micro-organisms and the substances they produce, that adheres to surfaces such as river beds or pipe interiors.

Biological fixation
With reference to nitrogen fixation: The process whereby atmospheric elemental nitrogen is reduced to ammonia, and which is carried out in the living world only by some free-living bacteria and cyanobacteria (blue-green algae) and by a few groups of bacteria in symbiotic association with plants.Lawrence (1995)

Biomagnification
The increasing concentration of a compound in the tissue of organisms as the compound passes along the food chain; resulting from the accumulation of the compound at each trophic level prior to its consumption by organisms at the next trophic level. Begon, Harper & Townsend (1986)

Biomass
The total mass or weight of all the living organisms in a given area or population.

Biomes
A broad, regional type of ecosystem characterised by distinctive climate and soil conditions and a biological community which is adapted to those conditions.

Biosphere
The zone of air, land, and water (both in lakes and in the sea) at the surface of the earth that is occupied by organisms.

Biotic
Applied to the living components of the biosphere or of an ecosystem, as distinct from the non-living, abiotic, physical and chemical components. Allaby and Allaby (1990)

Blue Green Algae
Common name for cyanobacteria. (See Cyanobacteria). Lawrence (1995)

Boreal forests (Taiga)
A broad band of mixed deciduous and coniferous woodland that stretches across most of northern North America, and Eurasia. Boreal forest is also known as Taiga.

Carbohydrates
Compounds of carbon, oxygen and hydrogen, of general formula Cx(H2O)y, including sugars (monosaccharides and disaccharides) and their derivatives, and polysaccharides such as starch and cellulose.
Lawrence (1995)

Carbon
The basic element of organic life which, together with hydrogen and oxygen, forms the fundamental molecules involved in organic structures.

Carbon dioxide
A colourless, odourless gas at normal temperatures, composed of one atom of carbon and two of oxygen. It makes up about 0.03% of the atmosphere by weight. Carbon dioxide is consumed in photosynthesis by green plants and produced by the respiration of plants and animals and the burning of fossil fuel.

Carbonate
A mineral containing the chemical group CO3 e.g. calcite (CaCO3). Carbonates are associated mainly with limestone and dolomite. Allaby and Allaby (1990)

Carbonic acid
A very weak acid formed from carbon dioxide and water. Sharp (1990)

Carnivores
Animals that feed on other animals, especially the flesh-eating mammals (Carnivora) such as dogs, cats, bears and seals, which feed on meat or fish. Lawrence (1995)

Chaparral
The scrub of west California and adjacent regions, in which the leaves of the trees and shrubs are evergreen, hard, thick, leathery and usually small. These adaptations allow the plants to survive the pronounced hot, dry season. Much of the chaparral has been derived by disturbance, principally burning of an earlier forest cover. Allaby (1994)

Chlorophyll
The green pigment in plants that functions in photosynthesis by absorbing radiant energy from the Sun, predominantly from blue (435-438nm) and red (670-680nm) regions of the spectrum. Allaby (1994)

Chloroplasts
The inclusion of (plastids) within plant cells which contain chlorophyll.Begon, Harper & Townsend (1986)

Class
Taxonomic group into which a phylum or a division is divided, and which is itself divided into orders. Lawrence (1995)

Climatic climax
A relatively stable, long-lasting ecosystem which is established after a successional series, usually best suited to the climate of the area.

Climax
A relatively stable, long-lasting community reached in a successional series; usually determined by climate and soil type.

Community
A general term applied to any grouping of populations of different organisms found living together in a particular environment; essentially, the biotic component of an ecosystem. Allaby (1994)

Colonisation
The successful establishment of an invading species in a habitat.Allaby (1994)

Cyanobacteria
A large and varied group of photosynthesizing bacteria which were formerly regarded as algae - hence the common name ‘blue green algae’. They are believed to have been the first oxygen-producing organism. Many species can fix atmospheric nitrogen. Some produce toxins which can become a health hazard when algal blooms occur. Lawrence (1995) Allaby (1994)

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Decomposer
Organism such as a maggot or bacteria that takes part in the decay of other organisms (plants and animals) by feeding on their remains.

Decomposition
The breakdown of complex organic material, undertaken by bacteria and fungi. Leads to the production of humus.

Deforestation
The removal of forests. Deforestation is often the result of human activities such as logging or farming and may lead to desertification.

Demes
A spatially discrete interbreeding of organisms with definable genetic or cytological characters (i.e. a subpopulation of a species). Allaby (1994)

Denitrification
Conversion of 'agriculturally useful' forms of nitrogen e.g. NO3-, NH4+ to dinitrogen (N2) gas.

Deserts
Areas receiving less than 25cm of rain a year. The term is more generally applied to any arid area. Deserts do not necessarily have a hot climate: cold deserts do occur.

Differentiation
The increasing specialization of the organization of the different parts of an embryo, as a multicellular organism develops from the undifferentiated fertilized egg. Lawrence (1995)

Diffusion
The movement of a substance down a concentration gradient from high concentration of the substance to low concentration, normally within gaseous fluids. Diffusion is caused by random movement of the particles of the substance so that ultimately it will lead to a homogenous distribution of the substance in the fluid.

Direct interference
The addition or removal of a material or element from an ecosystem by a plant which does not directly compete for it.

Diurnal
Daily; related to or performed in or lasting a day. Chambers (1994)

DNA
Deoxyribose nucleic acid (DNA) is found in the nuclei of cells and carries the necessary hereditary information to enable highly specific proteins to be constructed. Sharpe (1990)

Domain
A taxonomic rank introduced by Carl Woese in 1990 as the primary division of living organisms.

Dynamic equilibrium
A state where, although the mean state is constant, the forces acting continue to change and flux.In chemical terms dynamic equilibrium refers to the state in which there is no change in composition, unless there is a change in an external property.(e.g. temperature or pressure). This state only exists where the downward and reverse reactions are exactly balanced.

Ecosphere
See Biosphere.

Ecosystem
A specific biological community and its physical environment interacting in an exchange of matter and energy.

Edaphic climax
The climax vegetation that occurs as a result of soil induced factors.

Elements
A substance which cannot be further divided by chemical methods. The basic substances which build up chemical compounds. Sharpe (1990)

Environmental gradient
A continuum representing an factor such as mean annual precipitation, soil fertility, growing season length, etc. Each species will have a point on the continuum at which it experiences optimum growing conditions, and a range of tolerance representing the conditions in which it is able to survive.

Environmental spheres
See Atmosphere, Biosphere, Hydrosphere, Lithosphere, Mesosphere, Photosphere.

Enzymes
Molecules, usually proteins or nucleic acids, that act as catalysts in biochemical reactions.

Epiphytic plants
Plants that live on the surface of other plants but do not derive water or nourishment from them. Lawrence (1995)

Equilibrium
A state of even balance. Chambers (1994)

Erosion
The process of wearing away the land surface, by processes which transport rock debris. Erosion frequently refers to the loss of soil due to wind or as a result of washing away by water.

Essential elements
With reference to nutrition: elements necessary for the survival of the organism.

Eukaryotes
Organisms with cells possessing a membrane-bound nucleus in which the DNA is complexed with histones and organised into chromosomes, i.e. protozoans, algae, fungi, plants and animals. Begon, Harper & Townsend (1986)

Evolution
The development of new types of living organisms from pre-existing types by the accumulation of genetic differences over long periods of time. Lawrence (1995)

Extinction
The complete disappearance of a species from the earth.Lawrence (1995)

Facilitation model
This theory was put forward by Connell and Slatyer in 1977 but it is based upon the 'classic' theory of succession. It suggests that species that are being replaced in a succession will have altered their environment so that it is less suitable for their own needs whilst becoming more suitable for the needs of the replacing species. For example, as a plant species develops it changes the microclimate and soil around it due to both an increase in height, which produces calm areas, and organic decomposition, which will increase the fertility of the soil. Since this initial species may have been a pioneer species especially tolerant of harsh environments, it may not subsequently be able to survive competitively in the richer conditions it has helped to create.

Family
Taxonomic group of related genera, related families being grouped into orders. Familial names usually end in -aceae in plants and -idae in animals. Lawrence (1995)

Food Chain
A linked feeding series in an ecosystem: the sequence of organisms through which energy and materials are transferred, in the form of food, from one trophic level to another.

Food conversion ratios
The percentage of consumed food converted to body weight.

Food web
A complex interlocking series of individual food chains. (See: Food chain).

Fossil fuel
A fuel derived from the remains of ancient plant or animal life. The main forms used are oil, gas and coal.

Fundamental niche
The full range of conditions across which a species could survive and reproduce, in the absence of competition from other species.

Gametes
Reproductive cells produced by sexually reproducing organisms which fuse with another gamete of opposite sex or mating type to produce a zygote. Lawrence (1995)

Genetic information
The information for synthesizing RNA and protein, which is contained in DNA and RNA. Lawrence (1995)

Genera
The plural of genus. Taxonomic group of closely related species, similar and related genera being grouped into families. Generic names are italicized in the scientific literature e.g. Homo (man), Canis (wolves and dogs). Lawrence (1995)

Genes
The basic unit of inheritance, by which hereditary characteristics are transmitted from parent to offspring. At the molecular level a single gene consists of a length of DNA (or in some viruses RNA.) Each living cell carries a full complement of the genes typical of the species. Lawrence (1995)

Genus
One of the groups used in classifying organisms. Consists of a number of similar species, and occasionally of one species only. Similar genera are grouped in a family. Abercrombie, Hickman and Johnson (1980)

Global carbon budget
The carbon available to global systems, and its distribution within those systems.

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Halosere
A sere which is developed under saline conditions.

Hardpan
Hardened soil horizon, usually found in the middle or lower parts of the profile. Allaby and Allaby (1990)

Herbivores
An animal that feeds exclusively on plants. Lawrence (1995)

Higher plant
A plant of relatively greater complexity than the more simple, lower plants. Higher plants contain a vascular system of transport vessels including xylem. E.g. flowering plants, conifers, ferns.

Horizons
With reference to soils: A horizontal layer that can be distinguished from the layers below and above it. Identified by a coding system using a capital letter, sometimes followed by a subscript, such layers are used to diagnose soil types. Allaby and Allaby (1990)

Humus
Sticky, brown insoluble residue from the bodies of dead plants and animals. Humus gives soil its structure, coating mineral particles and holding them together. It also serves as a major source of plant nutrients.

Hydrological cycle
The circulation of water - in liquid, solid, or vapour form - through a series of reservoirs in the environment.

Hydrosere
A sere developed under (non-saline) wet conditions, such as lake margins.

Hydrosphere
The total body of water of the earth, i.e. the oceans, rivers, lakes, underground and atmospheric water.

Inhibitory effect
The ability to restrict the growth or potential for growth of competitors; an effect caused by some organisms in competition for nutrients.

Initial Floristic Competition theory
A succession theory which suggests that development at any one site depends largely on who gets there first: there is not necessarily an orderly progression because existing species will try to maintain their hold on the site. (Egler 1954). It is also known as the 'Inhibition Model' (Connell and Slayter 1977).

Inorganic
With reference to chemistry: of, or relating to, compounds which do not contain carbon. Also: not of animal or vegetable origin. Chambers (1994)

Ions
Atoms or molecules that have acquired an electric charge by the loss (cation; positive charge) or gain (anion; negative charge) of one or more electrons. Allaby and Allaby (1990)

Keystone species
Species that have a key role in an ecosystem, affecting many other species, and whose removal leads to a series of extinctions within the system. Lawrence (1995)

Kingdom
With reference to taxonomy: the primary division of the living world, until the category Domain was introduced above the level of Kingdom in 1990. Kingdoms are divided into phyla, for animals, and divisions for plants and other organisms. Lawrence (1995)

Leaching
The removal and transport of elements in solution from and through a soil.

Lichens
A composite organism formed from the symbiotic association of certain fungi and a green alga (or cyanobacterium), forming a simple thallus and found encrusting rocks, tree trunks etc., often in extreme environmental conditions. Lawrence (1995)

Life cycle
A series of developmental changes undergone by the individuals comprising a population, including fertilization, reproduction and the death of those individuals. Allaby (1994)

Limiting nutrient
One of a number of nutrients essential for plant or algal growth, the limited supply of which causes a halt to further growth as it is depleted.

Lithosere
A sere which develops on rock.

Lithosphere
The rigid uppermost portion of the mantle plus the overlying crust of the earth comprising surface rocks.

Litter
Accumulation of dead plant remains on the soil surface. Allaby and Allaby (1990)

Litter layer
See Litter.

Lower plant
A plant of relatively little complexity.

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Macronutrients
Nutrients required in large amounts by plants for growth: carbon, hydrogen, oxygen, nitrogen, phosphorus, sulfur, potassium, calcium and magnesium.

Mariculture
Mariculture is the growth of marine organisms (most commonly fish) in captivity for food or other uses.

Marine
Of, in, near or concerned with the sea. Chambers (1994)

Mediterranean
Biome that occurs around latitude 35oN and 35oS and is associated with warm temperate west coasts. Summers generally are hot and dry, winters mild, cool and rainy. The climate is strongly influenced by westerly air streams in winter and subtropical high pressure in summer. Allaby (1994)

Megafauna
Animals that are large enough to be seen with the naked eye. Allaby (1994)

Meiosis
A type of nuclear division which results in daughter nuclei each containing half the number of chromosomes of the parent. Lawrence (1990)

Mesosphere
Upper-atmospheric layer above the stratopause (at 50 km) through which temperature decreases with height up to about 80 km, where temperature reaches a minimum of about -90oC. Allaby and Allaby (1990)

Metabolism
The total of all the chemical reactions that occur within a living organism.Allaby (1994)

Micro fauna
Animals less than 200um long, such as protozoa, only visible under the microscope. Lawrence (1995)

Microclimate
The climate acting in a small area; a divergence through localised conditions from the average climate for a region.

Microhabitat
The habitat developed in a small localised area which is adapted to that site and situation; it may diverge markedly from the overall ecosystem of the region.

Micronutrient
Elements required in small amounts by plants for growth. Elements are: Fe, Na, Mn, Cu, Zn, Mo, B, Cl.

Microorganism
Any microscopic organism, including bacteria, viruses, unicellular algae and protozoans, and microscopic fungi (yeasts and moulds). Lawrence (1995)

Minerals
Naturally occurring homogenous inorganic substances of definite composition and crystal structure.

Mitochondria
An energy-producing body, threadlike to spherical shape, present in cytoplasm. Chambers (1994)

Mitosis
The normal process of nuclear division by which two daughter nuclei are produced, each identical to the parent nucleus, resulting in two daughter cells with identical nuclear contents. Allaby (1994)

Molar ratio
The ratio of each element within a molecule. For example, carbon dioxide (CO2) has a molar ratio of 1:2.

Molecule
A molecule is composed of two or more bonded atoms. Molecules are converted into other molecules during chemical reactions by breaking chemical bonds between atoms: new chemical bonds between different atoms are usually formed to create the product of a chemical reaction.

Monoclimax theory
The theory of monoclimax suggests that although there may be several seres developing within a climatic region the ultimate result will be convergence to a single climax form of vegetation, providing enough time is given and the process is free from interference.

Mutations
A change in the structure or amount of the genetic material of an organism.Allaby (1994)

Mutualistic interaction
A relationship between individuals of two different species in which both species benefit from the association

Niches
The functional position of an organism in its environment, comprising the habitat in which it lives, the period of time during which it occurs and is active there, and the resources it obtains there. Allaby (1994)

Nitrification
The oxidation of ammonium ion to nitrate, and the oxidation of nitrite to nitrate, carried out chiefly by a few groups of soil bacteria (nitrifiers) and also by a few species of fungi. Lawrence (1995)

Nitrogen cycle
The sum total of processes by which nitrogen circulates between the atmosphere and the biosphere or any subsidiary cycles within this overall process. Lawrence (1995)

Nucleic acids
A molecule that is composed of strings of NUCLEOTIDES forming a POLYNUCLEOTIDE CHAIN. Nucleic acids act as the genetic material of cells and occur as either DNA (two chains) or RNA (one chain). Hale & Magham (1988)

Nutrient cycles
The exchanges of elements between the living and non-living components of an ecosystem. Lawrence (1995)

Nutrients
Any substance that organisms take in and use for growth and maintenance.

Obligate predator
An predator that is very specific in its prey.

Omnivores
Animals that eat both plant and animal food. Lawrence (1995)

Order
Classification of organisms. Consists of a number of similar families. Similar orders are grouped into classes. Abercrombie, Hickman & Johnson (1980)

Organic matter
Substances formed by living organisms, often the product of their decay.

Oxalic acid (COOH2)
Also called ethanedioic acid (COOH2). An organic acid which is a by-product of the bacterial decomposition of organic matter.

Oxidised
To combine with oxygen. Also means to lose electrons. Chambers (1994)

Oxygen (O)
A gas without taste, colour or smell, forming part of the air, water, etc., supporting most forms of life and also necessary for combustion. (Atomic number 8; symbol O). Chambers (1994)

Parasite
An organism that lives in, or on, another living organism and derives subsistence from it, without rendering it any service in return. Chambers (1994)

Perennial
A plant that lives more than two years. Chambers (1994)

Permafrost
The permafrost is permanently frozen ground which extends 2000-3000 km south of the North Pole. It may be divided into three classes:i) Continuous, in which the subsoil never thaws. ii) Discontinuous, where patches of unfrozen ground occur. iii) Sporadic, where the area of unfrozen ground dominates. Seasonally areas of permafrost may thaw in their upper few centimetres allowing some plant growth.

Photosynthesis
The process by which green plants convert light energy into chemical energy (food), through the use of chlorophyll. The main chemical components are water and carbon dioxide.

Phylum
Classification of a primary group consisting of animals constructed on a similar general plan, and thought to be evolutionarily related. Lawrence (1995)

Physiographic climax
The climax vegetation that occurs as a result of terrain induced factors.

Phytographic climax
The climax vegetation that occurs as a result of light induced factors.

Phytoplankton
The small plants such as diatoms which float passively in seas and lakes. They form the base of many marine food chains.

Photosphere
The visible surface of the sun. The average temperature of the photosphere is 6000K.

Pioneer species
First species, usually mosses, lichens and micro-organisms that colonize a bare site as the first stage in a primary succession. Lawrence (1995)

Plagioclimax
A plant community arrested from seral development as a result of human interference.

Plasmalemma
The membrane round the outside of a living cell.

Polyclimax Theory
A theory that other factors are as important as climate in governing the climax vegetation. In any particular climatic region seres will develop a set of climax vegetations because of these other factors.

Population
A group of individuals of a species living in a certain area. Lawrence (1995)

Prairies
A temperate grassland of northern America dominated by xeromorphic grasses, which fall into three groups based on stature (tall, mid, and short) with a progressive decrease in rainfall. Allaby (1994)

Precipitation
With reference to the water cycle: Rain, hail, snow etc. Chambers (1994)

Predator
A predator is any species which uses another species, the prey, as a source of food. This ranges from algae feeding on bacteria through herbivores feeding on food plants to carnivores feeding on herbivores. The difference between a predator and a parasite is that a parasite can only survive if its host survives, whereas a predator eats and thereby kills its prey.

Prey
The creature or creatures that a predator hunts and kills as food. Chambers (1994)

Primary Producer
An organism that synthesises food molecules from inorganic compounds by using an external energy source: most primary producers are photosynthetic.

Primary productivity
The rate at which biomass is produced by photosynthetic and chemosynthetic autotrophs (mainly green plants) in the form of organic substances. Allaby (1994)

Primary succession
The sequence of development of vegetation on land that has not previously been influenced by vegetation (e.g. following lava flows, glacial retreat, newly exposed rock surfaces, etc). 

Proteins
One of the chief constituents of living matter, any one of a group of large organic molecules containing chiefly, carbon, hydrogen, oxygen, nitrogen and sulfur, and consisting of amino acids joined together by peptide bonds. Lawrence (1995)

Protozoans
Any member of the phylum Protozoa, including unicellular or acellular organisms. Hale & Margham (1988)

Productivity
The rate or efficiency of production of a given thing e.g. biomass.Chambers (1994)

Psammosere
A sere developed on sands or dunes.

Psyllid
An insect of the genus Psylla, sap-sucking plant pests. Chambers (1994)

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Rain forests
Forest biomes that develop in areas with an annual rainfall of more than 254cm. Lawrence (1995)

Realised niche
The range of conditions across which a species can be found. This is smaller than the fundamental niche due to competitive interactions with other species.

Relay Floristic Theory
See: Facilitation Model.

Respiration
The breakdown of chemicals (food) through oxidation to release energy for life. The main components of the reaction are water, oxygen and sugars. Note that nearly all living organisms respire, plants as well as animals.

RNA
Ribonucleic acid, large linear molecule of varied composition, made up of a single chain of a ribonucleotide subunits. Lawrence (1995)

Saline
Containing salt; a salt solution. Chambers (1994)

Savanna
Savanna or savannah. Tropical grassland areas situated between the humid equatorial regions and the arid zone in the mid-latitudes.

Scree slope
A slope on which an accumulation of coarse rock debris, added to by the weathering and release of fragments e.g. from a cliff face. Allaby and Allaby (1990)

Secondary climax
The climax vegetation resulting from the re-colonisation of a partially devastated climax vegetation.

Secondary consumers
A carnivore that preys upon herbivores. Allaby (1994)

Secondary sere
Second stage occurring in plant succession, initiated by the disruption of a previously existing seral or climax community by some major environmental disturbance and leading to a marked change in the stable vegetation community e.g. fire. Allaby (1990)

Secondary succession
The succession of seres following the partial devastation of an area.

Sere
A sere is the sequential development of a plant community or group of plant communities on the same site over a period of time. Succession at any particular site will progress through a sequence of seral stages.

Specialisation
The development of adaptations to a specific environment.

Species
A population of morphologically similar organisms that can reproduce sexually among themselves but cannot produce fertile offspring when mated with other organisms.

Stability
Ability of a community or ecosystem to withstand or recover from changes or stresses imposed from outside. Lawrence (1995)

Standing crop
Standing crop is the biomass present at any one time in any specific ecological component.

Steady state
The condition where the inputs and outputs (fluxes) of a reservoir balance one another.

Steppes
Dry, grassy, treeless, uncultivated and sometimes salt plain, as in central Europe and in Asia. Chambers (1994)

Stomata
Openings in epidermis of aerial part of plants, especially on undersides of leaves, through which air and water vapour enters the intercellular spaces, and through which carbon dioxide and water vapour from respiration is released. Lawrence (1995)

Stone age
A stage of culture before the general use of metal, divided into the Old Stone Age (Palaeolithic) and the New (Neolithic). Chambers (1994)

Stress
A physiological condition, usually affecting behaviour, produced by excessive environmental or psychological pressures. Allaby (1990)

Subsere
A secondary sere which develops after partial devastation of a site.

Substrates
A substratum; a base, a medium on which an organism can grow. Chambers (1994)

Succession
Succession refers to the sequential development of changes within a plant community. This is a natural process in which the species found in a given area change conditions to make the area less suitable for themselves and more suitable for others. This continues until the climax vegetation for an area dominates (See: Facilitation Model).

Symbiotic
Two organisms of different species living together in a persistent, close, and usually obligatory, association, not necessarily to their mutual benefit. Often used to describe an association in which both partners benefit, which is more properly called mutualism. Lawrence (1995)

System longevity
The ability of an ecosystem to survive wide shifts in the environment due to its stability and diversity.

Systematics
The study of the identification, taxonomy and nomenclature of organisms, including the classification of living things with regard to their natural relationships, and the study of variation and the evolution of taxa.Lawrence (1995)

Taxonomy
The science of classification, especially of living organisms, which arranges organisms into hierarchical groupings. Lawrence (1995)

Temperate Forest
Deciduous summer forest dominated by broad-leafed hardwoods, which occurs over large tracts in the mid-latitudes of Europe, North America and eastern Asia, but which is restricted in the Southern hemisphere to parts of Chilean Patagonia. Allaby (1994)

Temperate Grassland
A type of vegetation which includes in the northern hemisphere, the North American prairies and the Eurasion Steppe; and in the southern hemisphere, the veldt of South Africa, the pampas of Argentina, and the Canterbury Plains of New Zealand. Allaby (1994)

Temperate regions
The parts of the earth of moderate temperature, between the tropics and the polar circles. Chambers (1994)

Thunderstorm
A storm of fairly local scale in which strongly developed cumulonimbus cloud produces thunder and lightning, usually with rain and strong, gusting wind, and often with hail. Allaby and Allaby (1990)

Tolerance model
A succession theory proposed by Connell and Slatyer (1977) which suggests that replacements in a succession are not necessarily affected by earlier colonisers, and later species tend to be those which can tolerate lower levels of resources than earlier species.

Topography
The detailed study, description or feature of a limited area. With reference to land or the ocean bottom: a description of relief. Chambers (1994)

Trace elements
Elements which are present, or needed, in extremely small concentrations.

Transpiration
The evaporation of water from stomata in the aerial parts of plants, especially in the leaves. This evaporation cools plants and creates a pressure gradient which helps to enable the mass flow of water and dissolved nutrients upwards from the roots of the plant.

Trophic level
A step in the movement of energy through an ecosystem: an organism's feeding status in an ecosystem.

Tropical Grasslands
See Savanna

Tropical rain forests
Tropical rain forest is a species-rich biome consisting of stratified communities of broad leaved shrubs, trees, epiphytes, lianas, numerous insects, birds and other animals. Average monthly temperatures are normally in excess of 25oC and the range of temperature variation is small. Rainfall is abundant: between 1750 and 2500 mm per year, depending on the topography.

Tundra
Tundra is a treeless arctic or alpine biome where permafrost, low temperatures, high precipitation or wind have prohibited natural forest vegetation. Tundra may also be created when man, fire or animals destroy the natural forest and it fails to regenerate. High tundra is characterised by ground which is bare except for sparse lichen and grasses. Low tundra is characterised by a thick mat of vegetation often saturated with water and dotted with small ponds.

Veldt
Open unforested, or thinly forested grass-country. Chambers (1994)

Viruses
A protein-coated particle of RNA or DNA, capable of increasing rapidly inside a living cell. Chambers (1994)

Water column
A term used to describe the cross section of a body of water, with particular reference to the different processes taking place at various depths, e.g. the photic zone.

Water cycle
The cycle in which water from the sea evaporates into the atmosphere, later condenses and falls to the Earth as rain or snow, then evaporates directly back into the atmosphere, or returns to the sea by rivers. Chambers (1994)

Weathering
The process by which rocks are broken down and decomposed by the action of external agencies such as wind, rain, temperature changes, plants, and bacteria. Note that the process acts in situ with no transportation taking place and is therefore different from erosion. Two forms of weathering, chemical and physical, can take place and both may act at the same time.

Xeroseres
A sere which is developed in a dry, arid environment.

Xylem
Transport vessels found in the stems of all higher plants. They transport water and dissolved nutrients upwards from the roots to the leaves.

Zooplankton
Non-photosynthetic organisms and small invertebrate animals which are microscopic in size and occupy the upper water layers in freshwater and marine ecosystems. They live by eating smaller plants and or animals.

Zygotes
Fertilized eggs. Lawrence (1995)

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